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Endosperm is a terminal organ for storage of proteins and carbohydrates.Since maize has abundant different endosperm mutants and well established genetic analysis systems, it is no doubt the best model for study of endosperm development in grass crops.Opaque 2 (O2) and P-box Binding Factor (PBF) are two functionally conserved storage-protein transcription factors in grass crops belonging to bZIP and Dof transcription factor families, respectively.