生物圈演化的开始及其对生物地球化学的影响

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生物圈一开始演化就对气圈、水圈和岩石圈的有关演化具有深远的生物地球化学影响。在沉积记录中保存地壳演化的这几个圈同时相互影响的反馈作用为恢复生物地质学提供了证据。但是对这些证据的解释常常很困难的。生物成因和原生成因两者都需要证明,提出的每个形态或生物化学化石要确定它的可靠范围。生物圈和有关的演化的建立,必须批判性地估计到地球化学和沉积学的异常。间接的证据认为产生氧的自养生物大约在38亿年前就有了,但自由氧大约20亿年前才开始积聚。空气中和溶液中的各种还原物质是氧的沉淀槽(Sinks),它们保持光解和生物成因的氧相当于原始厌氧和显微亲氧原核细胞所要求的水平。已经证明的最古老生物成因和无疑的原生显微结构是上湖地区近于或大于2亿年地层中原核细胞。生物氧调解的改善,碳的不断分泌,充满了氧的沉淀槽开始了大气圈中氧的聚集就导致接近或晚于20亿年前臭氧层的形成。其后果是条带状含铁建造的基本结束,红层的突然出现,防护氧的厌氧细胞内作用,预示真核细胞的来临。可能的真核细胞出现在13亿年老的加利福尼亚州岩石中。它们是大的单细胞和直径粗、有分枝和隔壁的丝状体。真核细胞演变的可能后果是大气中氧的增加,碳酸根和硫酸根离子的增加和性别的出现。减数分裂肯定在7亿年前就逐渐形成了,也可能在13亿年前,大概和细胞的有丝分裂同时。不管是什么时间,它结束了真核细胞遗传机制的发展并且予示着组织、器官的分异和生命的高等形式。这些又带来对沉积作用,成岩作用、成矿作用潜在的反馈作用。最初的后生动物出现在7亿年前。它们没有外壳,依靠氧的简单渗透。生物外壳大约6亿年前产生,当时氧的含量水平已有利于出现高级的呼吸系统。 The first evolution of the biosphere has profound biogeochemical effects on the evolution of the balloon, hydrosphere and lithosphere. The simultaneous interaction of these few cycles of preservation of crustal evolution in sedimentary records provides evidence for the restoration of biogeology. However, the interpretation of these evidences is often difficult. Both biological and native causes need to be demonstrated, and each proposed morphological or biochemical fossil is to determine its reliable range. The establishment of biosphere and related evolution must be critically estimated to geochemistry and sedimentology anomalies. Indirect evidence suggests that oxygen-producing autotrophs existed about 3.8 billion years ago, but free oxygen began to accumulate about 2 billion years ago. The various reducing species in air and in solution are oxygen sinks that maintain photolytic and biogenic oxygen at levels that are required for the original anaerobic and micro-proophilic prokaryotic cells. The oldest biological genesis and undoubted native microstructure that has been proven is a prokaryotic cell in the strata near or greater than 200 million years in the Upper Lake region. The improvement of bio-oxygen mediation, the continuous secretion of carbon, the oxygen-filled sedimentation tank started the accumulation of oxygen in the atmosphere and led to the formation of the ozone layer close to or later than 2 billion years ago. The consequences of the basic end of the strip of iron-containing construction, the sudden emergence of red layer, protective oxygen anaerobic intracellular effects, indicating the advent of eukaryotic cells. Possible eukaryotic cells appear in the 1.3 billion-year-old rock of California. They are large, single-celled and coarse-grained filaments with branches and septums. Possible consequences of the evolution of eukaryotic cells are the increase of oxygen in the atmosphere, the increase of carbonate and sulfate ions and the appearance of sex. Meiosis certainly formed gradually 700 million years ago, probably 1.3 billion years ago, probably at the same time as cell mitosis. It ends the development of eukaryotic cytogenetic machinery no matter what time it is, and indicates the higher forms of tissue, organ differentiation, and life. These bring potential feedback on sedimentation, diagenesis and mineralization. The first metazoan appeared 700 million years ago. They have no shell and rely on simple infiltration of oxygen. The biological shell was created about 600 million years ago, when levels of oxygen were already favorable to the emergence of the advanced respiratory system.
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