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使用本文作者提出的分子绝对进化速率的计算公式和EF-1α,EF-2,NADH1和CytB等蛋白质分子,对原生生物的一些主要类群:藻类、真菌和原生动物的物种分歧时间和演化关系进行了定量计算.结果显示,原核生物沿着藻类→真菌→原生动物的方向演化.它们的演化关系为:(1)红藻自13.31亿年前分化出后,于11.74亿年前分化出绿藻,绿藻于4.527亿年前分化出藓类,4.37亿年前分化出蕨类;其他物种如石松类、肝苔类、团藻和小球藻分别于4.185亿年前、4.457亿年前、5.202亿年前和5.239亿年前分化出;(2)黏菌和卵菌均属于真菌进化分枝,分别于10.456亿年前和9.032亿年前分化出;(3)原生动物进化分枝中于10.45亿年前分化出孢子虫等,8.122亿年前领鞭毛虫与多细胞动物分化.这些结果均与基于化石记录的文献报道相符.“,”The divergence times and evolutionary relationships of some major protoctist clades were resolved by equation for molecular absolute evolutionary rates and several protein molecules:EF-1α,EF-2,NADH1 and CytB.The results showed that protoctists evolved from the route as:algae→fungi→protozoan.Their respective evolutionary routes are:(1) After the seperation of red algae at 1.331 Ga ago,the green algae separated from it around 1.1744 Ga ago, Mosses seperated at 0.4527 Ga ago,ferns seperated at 0.437 Ga ago.The divergence times of club-mosses,liverworts, Chlamydomonadales and Chlorellales are 0.4185,0.4457,0.5202 and 0.5239 Ga ago,respectively;(2) both Slime molds and oomycetes belonge to Fungi evolutionary branch,which separated about 1.0456 and 0.9032 Ga ago;(3) In protozoan branch,Apicomplexa and Choanoflagellates separated about 1.045 and 0.8122 Ga ago,respectively.Our results agreed well with the references work based on fossil records,and provided a new way to precisely resolve the evolutionary relationships of all protoctists.