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In this study, molecular cloning and functional characterization of an apple MdMYB4 (GenBank accession number: MDP0000582174) gene was reported.Sequence alignment and phylogenetic analysis showed that MdMYB4 was highly similar with Arabidopsis AtMYB4 and contained a typical R2R3 domain;MdMYB4protein was localized in the nucleus of transformed onion epidermal cells andhad a certain transcriptional activation by yeast one-hybrid assay;Expression analysis demonstrated that MdMYB4 extensively expressed in different tissues of apple.